Bauman A., Dunshea G., Fong J., Chan I.Z.W., Todd P.A. & Hoey A. (2021). Fear effects and group size interact to shape herbivory on coral reefs. Functional Ecology, 35(9): 1987-1999.

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Chan I.Z.W.†,1, Ngan Z.C.1, Naing L., Lee Y., V Gowri & Monteiro A. (2021). Predation favours Bicyclus anynana butterflies with fewer forewing eyespots. Proceedings of the Royal Society B, 288: 20202840.

     Featured by various media outlets: Straits Times , NUS, Mirage News, Science Daily,

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 Article in the Straits Times (20 Jun 2021) featuring our research published in Proceedings of the Royal Society B above.


Chan I.Z.W., Wang Y.W. & Todd P.A. (2020). Facial band colour in the mangrove crab Parasesarma peninsulare Shahdadi, Ng & Schubart, 2018 plays a role in mate recognition. Estuarine, Coastal and Shelf Science, 248: 106721.

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Chan I.Z.W.†,1, Rafi F.Z.1 & Monteiro A. (2019). Interacting effects of eyespot number and ultraviolet reflectivity on predation risk in Bicyclus anynana (Lepidoptera: Nymphalidae). Journal of Insect Science, 19(6): 19.

Lim A.Y.H.1, Chan I.Z.W.†,1, Carrasco L.R. & Todd P.A. (2019). Aposematism in pink warty sea cucumbers: independent effects of chromatic and achromatic cues. Marine Ecology Progress Series, 631: 157-164.

     Featured by the journal

Chan I.Z.W., Stevens M. & Todd P.A. (2019). PAT-GEOM: a software package for the analysis of animal colour pattern.  Methods in Ecology and Evolution, 10(4): 591-600.

Chan I.Z.W., Chang J.J.M, Huang D. & Todd P.A. (2018). Colour pattern measurements successfully differentiate two cryptic Onchidiidae Rafinesque, 1815 species. Marine Biodiversity, 49(4): 1743-1750.

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Chan I.Z.W., Stevens M. & Todd P.A. (2016). Quantifying shell pattern and colour polymorphism in the button snail Umbonium vestiarium (Mollusca: Gastropoda: Trochacea) and comparing morph frequencies between two populations using the Mantel test. Raffles Bulletin of Zoology, Supplement No. 32: 22-32 .

Submitted/In Preparation

Todd P.A.1, Chan I.Z.W.1, Wang W.Y., Boh Z., Soh H.S., Huang H. & Yeo D.C.J. (in preparation). Nature’s headlamps: a unique light-focusing structure in Parasesarma de Man, 1895 mangrove crabs.

Manuscript on the evolution of colour in nymphalids.

† Corresponding author
1 Co-first authors